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Official Brochure of Agate Fossil Beds National Monument (NM) in Nebraska. Published by the National Park Service (NPS).
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Agate Fossil Beds
Agate Fossil Beds National Monument
Nebraska
National Park Service
U.S. Department of the Interior
PHOTOS NPS UNLESS OTHERWISE CREDITED
A
bout 19–20 million years ago drought struck the western
Nebraska plains. Deprived of food, hundreds of animals
died around a few shallow water holes. Over time their
skeletons were buried in the silt, fine sand, and volcanic ash carried by the wind and reworked by streams. An ancient waterhole
with hundreds of fossilized skeletons is preserved today in the
Niobrara River valley at Agate Fossil Beds National Monument.
The early 1900s discovery of this deposit and others nearby was
important to the developing science of paleontology. Study of
these fossils—continuing today—has helped answer questions
about the past. But what were the conditions that created this
drought and brought these animals together so long ago?
Millions of years before the drought, in the Age of Dinosaurs,
erosion from mountain ranges to the west formed the bed of a
shallow sea. About the time dinosaurs went extinct, the Rocky
Mountain ranges were forming, the sea receded, and tropical
lowlands occupied what is now the Great Plains.
North America’s climate became ever cooler and drier, and volcanic activity in the western United States produced enormous
amounts of ash blown eastward. Ash-mantled plains were home
to great herds of plant-eating mammals and their predators. As
in today’s east African savannas, the rich volcanic soils supported
grasses that, together with small trees and bushes along shallow
streams, fed grass- and leaf-eaters. Many animals that thrived
here depended on the moderate climate for their survival, and
their numbers expanded to the capacity of the food available.
In time the climate grew more arid. The Rocky Mountains kept
rising and blocked the flow of moisture-laden air from the west.
With less rain came plants that could survive with less water.
Droughts were common. Streams dried up and grasses withered.
Water-dependent animals congregated at water holes between
times of feeding on the dwindling plants. Large animals like the
rhinoceros and the chalicothere Moropus, a distant relative of the
horse, finally could not travel far enough to find fresh forage, so
they died in the shallow water of the few remaining ponds.
Hundreds and thousands of some species died, littering the area
in and around water holes with their remains. In time the rains
returned, the streams filled, and the process of burial began. Silt,
sand, and ash covered the remains, burying them under several
feet of wind- and stream-transported sediment.
Fossils at Agate Fossil Beds
Top: Fossil exhibits in
visitor center.
Fossil photographs are not in relative scale.
JOURNAL AND FOSSILS—UNIVERSITY OF
NEBRASKA STATE MUSEUM
DENVER MUSEUM OF NATURAL HISTORY
What Animals Roamed Here? While some animals whose fossil remains were found at Agate
Fossil Beds are now extinct, others are represented by a few modern relatives or descendants.
Palaeocastor had powerful clawed forelimbs
for digging and long, curved teeth like modern
beavers. Herds of Stenomylus, gazelle-camels
about two feet tall, grazed grasslands beside
the three-toed, pony-sized rhinoceroses Menoc-
eras. The most common mammal in the bonebed, Menoceras may have roamed these plains
in large herds. Only a few oreodonts, about the
size of a sheep, have been found here, and they
are most common in the carnivore dens nearby,
where they were the prey of beardogs.
Fossil remains of the ancestors of the modern
horse, Parahippus, also have been found in the
waterhole but are rare. Horses became extinct
in North America millions of years after the dieoff event at Agate, not to return until brought
back by the Spaniards. Moropus was quite fantastic. Related to both the horse and rhinoceros,
it was large, had back legs shorter than the
front, with great clawlike hooves. It probably
browsed leaves of bushes and small trees.
pigs than to carnivores. Tracks of this huge scavenger have been found in the waterhole mud. It
broke bones with its teeth (bite marks show on
chalicothere limb bones). Discoveries in the 1980s
included fossil remains of beardogs and other
carnivores and their dens—one of the few paleontological sites of this type in the world.
Another large animal, Dinohyus, was a giant
entelodont related more closely to cows and
Discovery of the Fossils Most of the land that is
now Agate Fossil Beds National Monument was
Exploring Agate Fossil Beds
Visitor Center and
Museum Stop here
for information, activity schedules, exhibits
on fossils and artifacts,
and a short movie.
Open daily except
Thanksgiving, December 25, and January 1.
A picnic area is nearby. Call ahead for
educational programs.
It leads to the fossil
ized corkscrew burrows
of the small beaver
Palaeocastor.
Interpretive Trails Two
trails lead to important
fossil discovery areas.
The 2.7-mile Fossil Hills
Trail leads to University and Carnegie hills,
where most of the
digging took place in
the early 1900s. The
one-mile Daemonelix
—Devil’s Corkscrew—
Trail is near the River
Road-NE 29 junction.
Stay Safe, Protect the
Park Use common
sense to prevent accidents. • Rattlesnakes
live here, but you are
unlikely to encounter
one. Please stay on
the trails and out of
high grass. • Federal
law protects all natural and cultural features in the park. Do
not remove any fossils,
once part of the Agate Springs Ranch owned by
James and Kate Cook. They bought the ranch
from her parents in 1887, a few years after they
had found “a beautifully petrified piece of the
shaft of some creatures leg bone.”
Erwin H. Barbour of the University of Nebraska,
in 1892, was the first scientist to examine the
strange Devil’s Corkscrews at Agate, later recognized as the fossilized burrows of Palaeocastor.
Left to right: E.H.
Barbour’s 1892 field
book with Daemonelix
sketches and notes
on the Moropus bone
he mistook for a sloth
claw; fossil Dinohyus
tooth neatly fits the
bite mark on a Moropus bone; Dinohyus
skull illustrated O.A.
Peterson’s 1906 article
on vertebrate fauna
of western Nebraska;
fossil slab of Menoceras
bones; Harold Cook at
Colorado Museum of
Natural History.
In August 1904, O.A. Peterson of the Carnegie
Museum discovered the great bonebed with
the help of Harold J. Cook (above), son of James
and Kate. Scientists from Yale University, the
American Museum of Natural History, and other
institutions also worked here, mostly between
1904 and 1923. The competition to find the best
bones sometimes grew spirited. The work of
these bone hunters formed many outstanding
collections in museums around the world.
The Cook Collection of American Indian Artifacts
hits. • No overnight
camping or parking is
permitted. • Open
fires are prohibited.
• For firearms information, check the
park website.
Accessibility We strive
to make our facilities,
services, and programs
accessible to all. For
information, ask at
the visitor center, call,
or visit our website.
More information
Agate Fossil Beds
National Monument
301 River Rd.
Harrison, NE 69346
308-665-4111
www.nps.gov/agfo
animals, rocks, plants,
or artifacts. Leave
things as you find
them for others to enjoy, too. • There is
some private land
in the park. Respect
owners’ rights and do
not trespass. • Keep
pets on a leash and on
the trails. They are not
allowed in the visitor
center. • Lightning
kills; seek shelter before a thunderstorm
Agate Fossil Beds
National Monument is
one of over 400 parks
in the National Park
System. To learn more,
visit www.nps.gov.
James H. Cook was a
frontiersman, hunter,
and scout before he
settled on the Niobra
ra River. Cook first
met Chief Red Cloud
in 1874 when Yale
University Professor
Othniel C. Marsh came
to western Nebraska
looking for fossils.
The Oglala Lakota
(Sioux) were suspicious
of Marsh, because
most white men they
knew were gold seekers. But “Captain”
Cook helped convince
Red Cloud and the
other Oglala that
Marsh was what he
said he was, a bone
hunter.
Over the years Cook
often helped the
Oglala and Cheyenne.
A steadfast friendship
grew up between the
Cook family and the
Indians, who brought
gifts and told them
stories about individual
items. The family’s
collection now belongs to the park, and
many items are displayed in the visitor
center.
Pictographs painted
on hides—one of a
buffalo hunt and one
of Custer’s Last Stand—
saddles, bows, shirts
(including one of Red
Cloud’s that he gave
to the Cooks, right),
moccasins, bags, war
clubs, pipes, and guns
make this collection
an outstanding representation of Plains
Indian culture. Cook’s
interest in fossils led
his son Harold to
become a paleontologist, publishing many
papers and taking
part in important scientific research.
Preserved and available to researchers
here at the park are
the Cook collection,
other natural history
specimens, the paleontological library of
Harold Cook, and fam
ily correspondence,
books, and papers
that span four generations.
IGPO:2019—407-308/82302 Last updated 2019
The Age of Mammals
Life in the Cenozoic Era—the Last 65 Million Years
From simple beginnings great numbers and varieties of life
forms have evolved and populated the Earth. For 140 million
years before the Cenozoic Era, dinosaurs held dominion over the
land. Mammals also existed, but they were small and not abundant. As the dinosaurs perished the mammals took center stage.
Even as mammals increased in numbers and diversity, so did
birds, reptiles, fish, insects, trees, grasses, and other life forms.
The fossil record gives us a fascinating glimpse into the Cenozoic
Era. Without fossils we would have little way of knowing that
ancient animals and plants were different from today’s. With
fossils we discover that an extraordinary procession of organisms lived in North America and around the world. Species
changed as the epochs of the Cenozoic Era passed. Those that
could tolerate the changes in the environment survived. Other
Paleocene
Eocene
The Paleocene Epoch
began after dinosaurs
became extinct. Mam
mals that had lived in
their shadows for millions of years eventually
evolved into a vast number of different forms to
fill these newly vacated
environmental niches.
Many forms of these
early mammals would
soon become extinct.
Others would survive to
evolve into other forms.
In the Eocene Epoch
mammals emerged as
the dominant land animals. They also took to
the air and the sea. The
increasing diversity of
mammals begun in the
Paleocene continued
at a rapid pace in the
Eocene. The many variations included some of
the earliest giant mammals. Some were successful, some not. The
Began 65 million years ago
species migrated or became extinct. The fossil record tells these
stories, but the study of fossil remains, paleontology, also raises
many questions: What types of environments did these plants
and animals live in? How did they adapt to climatic changes?
How did different groups of plants and animals interrelate?
How have they changed through time?
Fossils are studied in the context in which they were found and
as one element in a community of organisms. Every fossil can
serve as a key to unlock knowledge, so the National Park Service
is especially concerned with the protection of these keys as the
questions unfold. The Cenozoic Era continues today—see the
right side of the chart below—and scientists estimate that as
Began 55 million years ago
Oligocene
Miocene
The Oligocene Epoch
was a time of transition
between the earlier and
later Cenozoic Era. The
once warm and moist climate became cooler and
drier. Subtropical forests
gave way to more temperate forests.
The abundance of mammals peaked in the Mio
cene Epoch. The refine
ment in life forms that
marked this epoch saw
many animals and plants
develop features recog
nizable in some species
today. The forests and
Began 34 million years ago
fossil record reveals
many mammals quite
unlike anything seen
today. Increasingly, however, there were forest
plants, freshwater fish,
and insects much like
those seen today.
Bats, the only type of
mammal ever to develop
the power of active
flight, took to the air
more than 52 million
years ago.
Many freshwater fish lived
in North American lakes
during the Eocene Epoch.
Gars (bottom), herring
(middle), and sunfish (top)
are similar in appearance
to those Eocene fish.
Fossil Butte NM
Fossil Butte NM
Groves of giant redwood
trees once grew throughout western North Amer
ica. Changes in climate
were responsible for these
trees’ shrinking range.
Florissant Fossil Beds NM
The variety of other
animals and plants also
increased, and species
became more specialized. Although dinosaurs
were gone, birds continued to flourish, and reptiles lived on as turtles,
crocodiles, lizards, and
snakes.
Delicate bones of shorebirds, including frigate
birds, are preserved in the
fine-grained sediment of
Eocene lake deposits.
As the Paleocene began,
most mammals were tiny,
like this rodent-like multituberculate. With time mammals grew in size, number,
and diversity.
Late in the Oligocene,
savannas—grasslands
broken by scattered
woodlands—appeared.
These changes caused
mammals, insects, and
other animals to keep
trending toward specialization. Some adapted to
the diminishing forests
by becoming grazers.
Early types of mammals
continued to die out as
more modern groups—
dogs, cats, horses, pigs,
camels, and rodents—
rose to new prominence.
Fossil Butte NM
Butterflies and many other
insect groups co-evolved
throughout the Cenozoic
with the increasing variety of flowering plants.
These insects became
important agents of
pollination.
Florissant Fossil Beds NM
Palm trees and crocodilians
thrived in the subtropical
forests of the Paleocene
and much of the Eocene.
many as 30 million species of animals and plants now inhabit
the Earth. This is a mere fraction of all life forms that have ever
existed. Scientists now think that about 100 species will become
extinct every day, a rate accelerated by human actions. Pollution
of the air and water; destruction of forests, grasslands, and other
ecosystems; and other adverse changes to Earth’s environment
challenge life’s very ability to survive. “Looking back on the long
panorama of Cenozoic life,” Finnish scientist Björn Kurten has
said, “I think we ought to sense the richness and beauty of life
that is possible on this Earth of ours.” It is no longer enough to
plan for the next generation or two, Kurten suggests. We should
plan “for the geological time that is ahead. . . . It may stretch as
far into the future as time behind us extends into the past.”
The variety of flowering
plants exploded just be
fore, during, and after the
Eocene. They would populate the land with all sorts
of new species of trees,
shrubs, and smaller plants.
Cattails grew in the shallows of Eocene freshwater
lake edges.
Coryphodon had short,
stocky limbs and five-toed,
hoofed feet, closely resembling the tapir. Its brain was
very small. The males had
large tusks. Coryphodon
also lived on land not far
from the shores of Fossil
Lake.
Fossil Butte NM
(Park names following captions
indicate where fossils were
found.)
Began 23 million years ago
savannas persisted in some
parts of North America.
Treeless plains expanded
where cool, dry conditions
prevailed. Many mammals
adapted for prairie life by
becoming grazers, runners, or burrowers. Large
and small carnivores
evolved to prey on these
plains-dwellers. Great
intercontinental migrations took place throughout the Miocene, with
various animals entering
or leaving North America.
Moropus was a distant
relative of the horse and
one of the more puzzling
mammals. For many years
paleontologists thought
its feet had claws rather
than hooves.
Agate Fossil Beds NM
Wyoming. Unlike modern
tapirs, Heptodon had a
very small snout.
Living in Eocene forests,
the first horse-like animals were barely bigger
than today’s domestic cat.
Throughout the Cenozoic
Era their size increased.
Their legs became longer,
and their feet changed
from many-toed to singlehoofed for faster running.
Their teeth evolved from
being adapted for brows
ing to being adapted for
grazing. Just a few of the
species in the evolution-
Pleistocene
Most life forms of the
Pliocene Epoch would have
been recognizable to us
today. Many individual
species were different, but
distinguishing characteristics of various animal and
plant groups were present.
Evidence of wet meadows
and of dry, open grassland
environments has been
found in the Pliocene.
Toward the end of this
epoch grasslands spread
across much of North
America, brought on by
an ever cooler, ever drier
climate. Horses and other
hoofed mammals and the
powerful, intelligent predators that preyed on them
continued to prosper.
The Pleistocene Epoch
began with widespread
migrations of mammals
and ended with massive
extinctions. It was also
a time when glaciers re
peatedly covered much
of North America.
Agate Fossil Beds NM
Ekgmowechashala
marked the end of the
original primate lineage in
North America. A small
lemur-like primate, it may
have used large skin folds
to glide from tree to tree.
Its name means “little cat
man” in Lakota, which the
discoverer understood to
be their name for monkey.
Began 2 million years ago
Known evidence of
humans living in North
America dates to about
12,000 years ago. In this
relatively brief period we
have had a profound effect
on the plants and other
animals here. Do we have
a responsibility to try to
limit our effects on other
species, or are humans
simply a natural agent of
extinction?
Mammut was a type of
mastodon that migrated
to North America in the
Lacking other defenses,
some larger rodents, such
as the dry-land beaver
Palaeocastor, lived in colonies beneath the High
Plains of North America.
Their burrows remain as
trace fossils today.
Rhinos were varied and
abundant during most of
the Cenozoic Era. Around
the world they ranged in
size from the three-foot-tall
North American species
Menoceras (shown here) to
a giant Asian species, the
largest land mammal yet
found in the fossil record.
Agate Fossil Beds NM
John Day Fossil Beds NM
Ancient tapirs such as
Heptodon browsed near
the shores of Fossil Lake
in what is now western
Pliocene
Began 5 million years ago
Tsetse flies occur
today in tropical
Africa and as fossils in
the Florissant formation.
Fossil Butte NM
Fossil Butte NM
These national parks (NP)
and national monuments
(NM) feature fossils from
the Cenozoic Era: Agate
Fossil Beds NM, 301 River
Road, Harrison NE 693462734; Badlands NP, PO
Box 6, Interior, SD 577509700; Florissant Fossil Beds
NM, PO Box 185, Florissant,
CO 80816-0185; Fossil
Butte NM, PO Box 592,
Kemmerer, WY 831010592; Hagerman Fossil
Beds NM, PO Box 570,
Hagerman, ID 83332-0570;
and John Day Fossil Beds
NM, HCR 82 Box 126, Kimberly, OR 97848-9701.
Pliocene. In the early Pleis
tocene another elephant
group called mammoths
joined the mastodons. By
the late Pleistocene masto
dons and mammoths both
became extinct, possibly
because of climatic changes
or hunting by early people.
Endangered species today
include the loon (top),
timber wolf (middle), and
Kemp’s ridley sea turtle
(bottom). The National
Park Service is among the
many public agencies and
private organizations
entrusted with helping to
protect endangered plants
and animals and to preserve the diversity of life
throughout North America.
Hagerman Fossil Beds NM
Daphoenodon was carnivorous. It differed from
the earliest true dogs of
the Oligocene Epoch. Its
Florissant Fossil Beds NM
Fossil Butte NM
ary history of horses are
shown here in silhouette
across this chart. Fossil
horses occur at many sites
in the National Park
System.
so-called “beardog”
family eventually went
extinct.
Agate Fossil Beds NM
Willow, alder, birch, and
elm grew on the ancient
river plains of the Plio
cene. These same plants
grow along streams and
rivers today.
Hagerman Fossil Beds NM
Oreodonts, a group of
sheep-like animals, were
successful in the Eocene
and Oligocene. By the end
of the Miocene they had
completely died out.
Daeodon (formerly called
Dinohyus, “terrible hog”)
had bone-crushing teeth
enabling it to scavenge
the remains of other
grassland animals.
TREE ILLUSTRATION NPS/JOHN DAWSON. ALL
OTHER FULL-COLOR ILLUSTRATIONS NPS/KAREN
BARNES. SOME DEPICTIONS OF MAMMAL
SPECIES FOLLOW FOSSIL RECONSTRUCTIONS
AS REPRESENTED IN R.J.G. SAVAGE AND
M.R. LONG’S MAMMAL EVOLUTION: AN
ILLUSTRATED GUIDE, NEW YORK: FACTS ON
FILE PUBLICATIONS, 1986. THE DRAWINGS ARE
NOT TO SCALE.
Agate Fossil Beds NM
Horses such as this early
zebra-like version of the
modern horse were su
perbly adapted to life on
the grassy plains.
Badlands NP
Hagerman Fossil Beds NM
The tiny gazelle-camel
Stenomylus probably
grazed in herds for protection from predators.
Agate Fossil Beds NM


