The Kofa National Wildlife Refuge is located in Arizona in the southwestern United States, northeast of Yuma and southeast of Quartzsite. The refuge was established to protect desert bighorn sheep in the Sonoran Desert. Broad, gently sloping foothills as well as the sharp, needlepoint peaks of the Kofa Mountains are found in the rugged refuge. The small, widely scattered waterholes attract a surprising number of water birds for a desert area. A wide variety of plant life is also found throughout the refuge.
Native Pollinators at Kofa National Wildlife Refuge (NWR) in Arizona. Published by the U.S. Fish & Wildlife Service (USFWS).
Kofa NWR
https://www.fws.gov/refuge/kofa/
https://en.wikipedia.org/wiki/Kofa_National_Wildlife_Refuge
The Kofa National Wildlife Refuge is located in Arizona in the southwestern United States, northeast of Yuma and southeast of Quartzsite. The refuge was established to protect desert bighorn sheep in the Sonoran Desert. Broad, gently sloping foothills as well as the sharp, needlepoint peaks of the Kofa Mountains are found in the rugged refuge. The small, widely scattered waterholes attract a surprising number of water birds for a desert area. A wide variety of plant life is also found throughout the refuge.
U.S. Fish & Wildlife Service
Public Use and Hunting Regulations 2019-2020
Kofa National Wildlife Refuge
General Information
Kofa National Wildlife Refuge (NWR) was
established in 1939 and is managed by the United
States Fish and Wildlife Service. The refuge
encompasses 666,641 acres of Sonoran desert
habitat and is home to the desert bighorn sheep,
Sonoran pronghorn and the California fan palm.
Other notable wildlife species found in the area
include white-winged dove, desert tortoise, desert
mule deer, and desert kit fox. A wide variety of plant
life is found throughout the refuge.
Bighorn sheep are found primarily in the two
mountain ranges that dominate the refuge
landscape—the Kofa and Castle Dome Mountains.
Although these mountains are not especially high,
they are extremely rugged and rise sharply from
the surrounding desert plains, providing excellent
bighorn sheep habitat. Endangered Sonoran
pronghorn were reintroduced in 2011 to a semicaptive breeding pen on the refuge. The first
Sonoran pronghorn on Kofa were released into the
wild in 2013; they now roam free on Kofa NWR and
adjacent lands. The wild population has grown to
about 75 animals through successful reproduction
and additional releases.
Water is scarce in the desert. By improving natural
water holes and creating new ones, refuge managers
have increased the availability and reliability of
water for wildlife.
Many wildlife species have benefitted from these
water developments. A long-term average of
approximately 400-800 desert bighorn sheep live
on the refuge. Beginning in the 1950s, the refuge
has provided over 560 desert bighorn sheep
for transplanting to other parts of Arizona and
neighboring states.
Palm Canyon, at the west end of the Kofa
Mountains, is well known for its stand of California
fan palms, the only native palm species in Arizona.
Fewer than 100 trees remain in Palm Canyon; these
palms are probably remnants from an era when this
area was wetter and cooler.
In the late 19th and early 20th centuries, numerous
silver, gold and lead mines were established
throughout the refuge. The most notable was the
“King of Arizona Mine,” which produced 3.5 to 4.0
million dollars in gold between 1897 and 1910. The
Kofa Mountains are a contraction from the name of
the “King Of Arizona” gold mine.
Most refuge roads are not maintained. Many are
passable only by four-wheel drive vehicles. Some
roads may be passable by two-wheel drive, high
clearance vehicles (see map). Vehicles may be
damaged by brush or rocks, or may become stuck in
sandy areas. Private lands (inholdings) occur within
the refuge and may be posted. Before traveling on
the refuge, visitors should contact refuge visitor
center to learn about current road conditions. There
are no facilities on the refuge for towing, gasoline,
sanitation, or drinking water. Cellular phone
coverage is absent over most of the refuge. Visitors
are advised to let someone know where they going
and when they plan to return.
Public Use Regulations
The following is a summary of refuge regulations.
For additional detail or clarification, contact the
refuge visitor center. All public access, use or
recreational activity, unless specifically authorized,
is prohibited.
Wilderness
Passage of the 1990 Arizona Desert Wilderness
Act included 547,700 acres of the Kofa NWR.
Wilderness status acknowledges the relatively
pristine character of the refuge and dictates some
management restrictions. No vehicular travel,
including bicycles, is permitted into wilderness
areas. Only foot or horseback travel is permitted.
Vehicles
Vehicular traffic is limited to designated roads which
can be identified by numbered markers at junctions.
Off-road vehicle travel is strictly prohibited.
Vehicles may pull off and park only up to 100 feet
from designated roads. No vegetation or other
refuge features may be damaged. This includes all
motorized or wheeled vehicles, such as ATVs, UTVs,
quadratracs, motorcycles, bicycles etc. All motor
vehicle operators must be licensed and insured for
highway driving. Speed is limited to 25 MPH, or
less as posted.
Animal and Plant Life
Collecting, possessing, molesting, disturbing,
injuring, destroying, removing or transporting any
plant or animal or part thereof (alive or dead) is
prohibited (except for legally taken game).
Firearms and Explosives
Carrying, possessing, or discharging explosives
(including fireworks) on the refuge is prohibited.
Persons may possess, carry, and transport firearms,
in accordance with all Federal and state laws.
Unauthorized discharge of firearms or target
practice is prohibited. For additional information
see the Hunting Regulations section.
Weapons Other than Firearms
Use or possession of other weapons, such as
crossbows, bows and arrows, and airguns is only
permitted in conjunction with authorized and
permitted hunting activities.
Disposal of Waste
Littering and the dumping of sewage or liquid
wastes on the refuge are prohibited. Do not bury
trash—wildlife will di
U.S. Fish & Wildlife Service
Palm Canyon Trail
Kofa National Wildlife Refuge
Welcome to Kofa’s Palm Canyon Trail
The survival of these trees is directly
dependent on the microclimate in this
protected canyon. The palm trees are
only able to live in the narrow side
canyons where direct sunshine is limited
and moisture is available.
How to find Palm Canyon
Finding Palm Canyon is relatively easy.
Take Highway 95, running north and
south between Yuma and Quartzsite,
Arizona, to milepost 85. Approximately
63 miles north of Yuma or 18 miles south
of Quartzsite, watch for the brown Palm
Canyon sign which indicates the junction
with a dirt road. Follow the dirt road east
for seven miles to a parking area. The
road is passable in a passenger car, but
the ride may be rough.
Palm Canyon
All photographs USFWS.
Tucked away in the narrow, rugged
canyons of the Kofa Mountains is the
California fan palm, the only native
species of palm tree in Arizona. The
small, scattered clusters of palms in Palm
Canyon are a major visitor attraction on
Kofa National Wildlife Refuge. Hundreds
of visitors each year hike the Palm
Canyon Trail, a National Recreation
Trail, to see the palms which seem
unnatural in the desert landscape.
The California fan palms (Washingtonia
filifera) found in Palm Canyon are
probably descendants of palms growing
in this region during the last periods
of North American glaciation. Some
botanists theorize that the trees, which
are not date-bearing, gradually spread
into these canyons and other protected
niches as the climate warmed to desert
conditions. Other researchers have
suggested that the trees may have spread
from other palm groves by birds or
coyotes carrying seeds in their digestive
tracts. Since palm trees do not grow
annual rings like other trees, it is very
difficult to say how old the trees might be.
California fan palms typically live for 80
to 90 years.
The narrow canyon above the parking
lot is Palm Canyon. A half-mile foot trail
starts at the upper end of the parking
area and leads into the canyon. The trail
is easy to follow, but is rough most of the
way due to large rocks and some steeper
sections. Allow an hour to make the
round-trip.
Desert Vegetation
As you follow the trail across washes and
up hills, you will pass several species of
plants common to the Sonoran Desert.
The palo verde (Cercidium spp.), a
small tree with bright green branches
and stems, rarely has leaves as it sheds
them in times of drought. Its branches
and twigs have enough chlorophyll to
produce all the energy the tree requires.
Ironwood (Olneya tesota), a grey-green
tree with small leaves, is also present in
the canyon. Its leafy stems are covered
with thorns, as are the branches and
stems of many other desert plants.
Near the end of the trail, you will notice
numerous small bushes with holly-like
leaves. This plant is Kofa Mountain
barberry (Berberis harrisoniana). It
is found only in the southwest corner of
Arizona in the Kofa, Ajo and Sand Tank
Mountains. It is not very common, even
on the refuge.
Ironwood
Palo Verde
The Palm Canyon trail ends at a small
sign on a slightly elevated area near the
middle of the canyon. By looking up at
the narrow north-trending side canyon,
you can see the palms. For a short time at
mid-day, the trees are well-lit for taking
photographs. The rest of the time, the
trees are in shade.
In 2001, forty-one trees were counted in
the main grove. About half of those were
adult size with a trunk height of 20 feet or
more. Some smaller trees are becoming
established at the base of the larger
trees. There are a few additional palms
scattered throughout Palm Canyon and
in Fishtail Canyon, the next canyon to
the southeast.
As the fronds (large leaves) on the
California fan palm die, they fold down
around the trunk of the tree and form a
“petticoat.” The petticoat on younger trees
extends from the ground to the top of the
tree. The fronds on the older trees in Palm
Canyon do not form a lengthy petticoat
because they tend to self-prune. The fallen
fronds decay and form an organic layer in
which new trees germinate.
Fire
You will notice from the viewing point
that some of the tree trunks are black.
A fire burned through the grove in
1953, seriously damaging the trees.
Fortunately, most of the palms survived
and young trees have become established.
It is a very steep and difficult climb up
to the palm trees. If you decide to try it,
plan on an extra 30-40 minutes to get up
to the trees and back to the bottom of
the canyon.
Nolinas
As you look around the canyon, you will
see palm-like plants which can sometimes
be confused with the California fan palm.
These plants are Bigelow’s nolina or
Bigelow’s beargrass (Nolina bigelovii)
and are growing out of cracks and on
ledges, especially on the north wall of the
canyon. The nolinas are much smaller
than palms and do not develop a trunk.
Wildlife
You may see a variety of wildlife as you
walk along the trail. Keep watching
the skyline on both si
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Crystal Hill Area
Kofa National Wildlife Refuge
Crystal Hill, located in the northwest
corner of Kofa National Wildlife Refuge
in the Livingston Hills, is the only area
within the refuge where recreational rock
or mineral collecting is permitted. Quartz
crystals are hidden in the washes and on
the rocky slopes of Crystal Hill, making
this an ideal location for rockhounding
or collecting. While searching for quartz
crystals at the top of the hill, visitors can
also enjoy the 360 degree view of the
surrounding area.
Creation of the Mountains
The mountains of Kofa National Wildlife
Refuge, which rise abruptly from the flat
desert, are characteristic of the Basin
and Range Province which covers much
of the western United States and extends
into northwestern Mexico. In this
physiographic region, the Earth’s crust
and upper mantle thinned and cracked
as they were stretched and pulled apart.
Quartz Formation
Quartz, belonging to the class of silicates,
comprises approximately 12 percent of
the Earth’s crust. It is produced when
the two most abundant elements in the
surface of the Earth, oxygen and silicon,
bond. Quartz crystallizes when volcanic
magma cools and becomes solid.
A fault created by crustal movement.
Illustration from http://clasfaculty.ucdenver.edu/
callen/1202/Battle/Build/Faulting/Faulting.html
This crustal movement created faults
and eventually led to the creation of
mountains and valleys as the Earth’s
surface uplifted on the upthrown side
of the fault and down-dropped on the
lower side.
Over time, the block-faulted mountains
weathered and eroded. Volcanic episodes
also occurred within the refuge and thus
the mountain ranges of Kofa National
Wildlife Refuge contain old lava layers
and volcanic rocks, such as basalt,
volcanic tuff and obsidian.
Quartz appears in a variety of forms and
shapes and varies from being clear to
almost opaque. It may be found in the
form of large grains, crystals or veins.
Crystals can appear either as a single
point or in a cluster. Pure quartz is white
or colorless, while other types of quartz
may be rose, yellow, green, blue, violet,
brown, or black depending on the trace
elements they contain.
The Livingston Hills are known to have
both quartz veins and crystals. Some
veins in the hills contain tourmaline and
pyrite, two other minerals. Most of the
quartz crystals found at Crystal Hill are
either clear or milky white.
Rockhounding and Collecting on the
Refuge
When searching for quartz crystals on
the refuge, it is important to remember
that recreational (noncommercial) rock
or mineral collecting is not permitted
beyond the boundaries of the Crystal
Hill Area (see map). Collection of rocks
is limited to 10 specimens or 10 pounds
(whichever occurs first) in any 12-month
period. Rock or mineral collection is
limited to materials that are exposed
and collectable without the use of tools,
including metal detectors. Digging
is prohibited. Please see the Kofa
National Wildlife Refuge leaflet Public
Use and Hunting Regulations for more
information about regulations.
For Further Information
U.S. Fish and Wildlife Service
Kofa National Wildlife Refuge
9300 East 28th Street
Yuma, AZ 85365
928/783-7861
Monday–Friday: 8:00 a.m.–4:30 p.m.
February 2014
Front, illustration of a desert bighorn.
Illustration by Elizabeth Montgomery
Desert bighorn at Kofa National Wildlife
Refuge. / USFWS.
U.S. Fish and Wildlife Service
Watchable Wildlife
Kofa National Wildlife Refuge
Welcome to wildlife watching on Kofa
National Wildlife Refuge
Kofa NWR encompasses 665,400 acres
of Sonoran Desert in southwestern
Arizona. Broad valleys, gently rolling
foothills, jagged peaks, and steep cliffs
are found within this rugged Refuge.
The Refuge receives little annual rainfall
and is characterized by warm winters
and scorching summer heat. Despite
these harsh conditions, a diverse group of
mammals, amphibians, and reptiles are
found on Kofa NWR.
Black-tailed jack rabbits, zebratailed lizards, western diamondback
rattlesnakes, and desert bighorn sheep
are several of the more commonly seen
animals on the Refuge. Other species,
including the coyote, Gila monster, desert
tortoise, and kit fox, may be spotted on
occasion, particularly at dawn or dusk.
Listed here are mammals, reptiles, and
amphibians recorded on Kofa NWR. Birds
are on a separate checklist. Additional
species may be present on the Refuge.
Symbols used in this list follow:
* Nonnative
† Native to southeast Arizona border area
Mammals
Order Chiroptera (Bats)
Family Molossidae
Western Mastiff-bat (Eumops perotis)
Pocketed Free-tailed Bat
(Nyctinomops femorosacca)
Mexican Free-tailed Bat
(Tadarida brasiliensis)
Family Phyllostomidae
California leaf-nosed bat
(Macrotus californicus)
Desert bighorn sheep. Susanna Henry / USFWS
Mule Deer. Katrina Krebs / USFWS
Family Vespertilionidae
Pallid Bat (Antrozous pallidus)
Townsend’s Big-eared Bat
(Corynorhinus townsendii )
Big Brown Bat (Eptesicus fuscus)
Spotted Bat (Euderma maculatum)
California Myotis (Myotis californicus)
Arizona Myotis (M. occultus)
Cave Myotis (M. velifer)
Yuma Myotis (M. yumanensis)
Canyon Bat
(Parastrellus hesperus)
Order Lagomorpha (Rabbits and Hares)
Family Leporidae
Black-tailed Jack Rabbit
(Lepus californicus)
Desert Cottontail (Sylvilagus audubonii)
Order Rodentia (Rodents)
Family Cricetidae
White-throated Woodrat
(Neotoma albigula)
Desert Woodrat (N. lepida)
Southern Grasshopper Mouse
(Onychomys torridus)
Brush Mouse (Peromyscus boylii)
Canyon Mouse (P. crinitus)
Cactus Mouse (P. eremicus)
Deer Mouse (P. maniculatus)
Western Harvest Mouse
(Reithrodontomys megalotis)
Family Erethizontidae
Porcupine (Erethizon dorsatum)
Family Geomyidae
Botta’s Pocket Gopher
(Thomomys bottae)
Family Heteromyidae
Bailey’s Pocket Mouse
(Chaetodipus baileyi)
Long-tailed Pocket Mouse (C. formosus)
Rock Pocket Mouse (C. intermedius)
Desert Pocket Mouse (C. penicillatus)
Desert Kangaroo Rat
(Dipodomys deserti)
Merriam’s Kangaroo Rat (D. merriami)
Arizona Pocket Mouse
(Perognathus amplus)
Little Pocket Mouse (P. longimembris)
Family Sciuridae
Harris’ Antelope Squirrel
(Ammospermophilus harrisii)
Round-tailed Ground Squirrel
(Xerospermophilus tereticaudus)
Order Soricomorpha (Shrews)
Family Soricidae
Desert Shrew (Notiosorex crawfordi)
Order Carnivora (Carnivores)
Family Canidae
Coyote (Canis latrans)
Gray Fox (Urocyon cinereoargenteus)
Kit Fox (Vulpes macrotis)
Family Ranidae
†Tarahumara Frogs (Rana tarahumarae)
Family Felidae
Bobcat (Lynx rufus)
Mountain Lion (Puma concolor)
Reptiles
Family Mephitidae
Striped Skunk (Mephitis mephitis)
Western Spotted Skunk
(Spilogale gracilis)
Family Mustelidae
Badger (Taxidea taxus)
Family Procyonidae
Ringtail (Bassariscus astutus)
Order Artiodactyla (Even-toed ungulates)
Family Antilocapridae
Sonoran Pronghorn
(Antilocapra americana sonoriensis)
Family Bovidae
Desert Bighorn Sheep
(Ovis canadensis Mexicana)
Family Cervidae
Mule Deer
(Odocoileus hemionus crooki)
Family Tayassuidae
Collared Peccary (Pecari tajacu)
Order Perissodactyla
Family Equidae
*Burro (Equus africanus asinus)
*Horse (E. caballus)
Amphibians
Order Anura (Toads and Frogs)
Family Bufonidae
Colorado River Toad (Bufo alvarius)
Great Plains Toad (B. cognatus)
Red-spotted Toad (B. punctatus)
Family Scaphiopodidae
Couch’s Spadefoot (Scaphiopus couchii)
Order Testudines (Turtles and Tortoises)
Family Testudinoidae
Desert Tortoise (Gopherus agassizii)
Order Squamata (Lizards and Snakes)
Family Crotaphytidae
Collared Lizard
(Crotaphytus insularis bicinctores)
Long-nosed Leopard Lizard
(Gambelia wislizenii wislizenii)
Family Gekkonidae
Western Banded Gecko
(Coleonyx variegatus variegates)
Family Helodermatidae
Banded Gila Monster
(Heloderma suspectum cinctum)
Family Iguanidae
Desert Iguana
(Dipsosaurus dorsalis dorsalis)
Chuckwalla (Sauromalus obesus obesus)
Family Phrynosomatidae
Zebra-tailed Lizard
(Callisaurus draconoides rhodostictus)
Desert Horned Lizard
(Phrynosoma platyrhinos calidiarum)
Desert Spiny Lizard
(Sceloporus magister magister)
Colorado Desert Fringe-toed Lizard
(Uma notata rufopunctata)
Long-tailed Brush Lizard
(Urosaurus graciosus graciosus)
Tree Lizard (U. ornatus symmetricus)
Side-blotched Lizard
(Uta stansburiana elegans)
Night Snake
(Hypsiglena torquata ochrorhyncha)
Common Kingsnake
(Lampropeltis getulus californiae)
Coachwhip
(Masticophis flagellum piceus)
Pine Gopher Snak
U.S. Fish & Wildlife Service
Native Pollinators
Kofa National Wildlife Refuge
When the desert ironwood tree blooms
in the spring, it begins to buzz with life.
Bees bustle around the tree, attracted to
its pink and white flowers. Bees, as well
as birds, bats, butterflies, beetles, flies,
wasps, and other insects, are important
pollinators. These animals facilitate
fertilization in plants like the desert
ironwood by transferring pollen between
flowers of the same species, allowing
the plant to produce successful seeds.
Pollinators are crucial to all national
wildlife refuges as between 75 and 95
percent of all flowering plants in the
world cannot self-pollinate.
Birds
Globally, over 2,000 species of birds feed
on nectar and eat the insects and spiders
associated with nectar-bearing flowers.
Birds have little or no sense of smell and
thus the flowers they visit are usually
odorless. These flowers tend to be rich in
nectar and have large, sticky pollen that
cling to the birds’ feathers.
When hiking on Kofa National Wildlife
Refuge, you might be startled by a loud
humming noise. The culprit is likely a
hummingbird (family Trochilidae), a
small and colorful pollinator with a long,
tapered bill and bristled tongue. The noise
results from the flapping of its wings
which beat up to 80 times per second. This
adaptation, as well as the hummingbird’s
ability to fly in any direction (including
upside down and backward), makes it an
excellent pollinator as it can hover in front
of flowers when feeding.
The hummingbird is generally attracted
to long, tubular, and brightly colored
flowers, though it will visit nearly any
nectar-producing flower. When the
hummingbird feeds on the flowers’ nectar,
sticky pollen attach to the bird’s bill or
feathers. As the hummingbird flies from
flower to flower, it spreads the pollen and
fertilizes the plant.
There are five species of hummingbirds
found on Kofa National Wildlife Refuge:
Costa’s hummingbird, black-chinned
hummingbird, Anna’s hummingbird,
broad-tailed hummingbird, and rufous
hummingbird.
Pollinators at Kofa National Wildlife Refuge. / Katrina Krebs, USFWS
The white-winged dove (Zenaida
asiatica), named for the white stripe at
the edge of its wing, is another common
bird pollinator on the refuge. In the
desert southwest, the white-winged
dove is considered a saguaro specialist.
It is almost exclusively dependent on
saguaros for nutrients and water during
the breeding season, eating the nectar,
pollen, fruits, and seeds of saguaros.
When visiting saguaro flowers, the whitewinged dove inadvertently pollinates
the cactus. It also helps disperse seeds
contained within the saguaro fruit it eats,
either through its feces or regurgitation
to its young. The white-winged dove is so
reliant on saguaros that its migration and
breeding season matches the flowering of
the saguaro.
Bees
Bees are the most diverse group of
pollinators with more than 1,000 species
in the southwestern United States and
over 4,000 species in North America.
Bees are generally attracted to flowers
that are blue or yellow, or a mixture of
these colors. They are unable to see the
color red, but can see ultraviolet light
unlike humans. Many flowers that attract
bees have ultraviolet patterns on the
petals. These patterns, called nectar
guides, help bees locate the center of the
flower so they can quickly find the nectar.
On Kofa National Wildlife Refuge, cactus
bees (Diadasia spp.) are an incredibly
important type of bee as they favor the
flowers of, unsurprisingly, cacti. The
cactus bee is fuzzy, brown and about the
size of a honey bee. It is a solitary ground
nester and is active primarily between
April and June when many of the cacti
are blooming on the Refuge.
The cactus bee crawls inside the flowers
of saguaros, prickly pears, barrel cacti,
teddybear chollas, and other cacti to feed
on their nectar, gathering pollen on its
hairy body all the while. As the bee flies
from flower to flower, the pollen is spread
and the flowers are fertilized. The cactus
bee also mixes some of the pollen with
nectar and stores it in its nest as food for
its young.
Butterflies
Butterflies are valuable pollinators on
the refuge, especially of brightly colored
flowers with sweet scents. Yet with their
longer legs, butterflies prove to be less
effective pollinators than bees as they do
not collect as much pollen on their bodies
while searching for nectar. They do serve
another purpose on the refuge though:
butterflies are an important food source
for other animals, particularly birds.
The marine blue (Leptotes marina) is
found year-round in the Sonoran Desert.
The upperside of its body is blue, while
the undersides of its wings are striped
brown and white. Host plants of this
butterfly species include legumes, such as
fairydusters, mesquite and acacias.
White-winged dove. / Katrina Krebs, USFWS
Wasps
A number of wasps that pollinate plants
resemble bees. However, they can
generally be distinguished from bees
by their narrow ab