The Bill Williams River National Wildlife Refuge protects the lower course of the Bill Williams River, to its mouth at Lake Havasu reservoir, in western Arizona. It is located within eastern La Paz and Mohave Counties, in the Lower Colorado River Valley region. Recreation activities include nature walks, bird watching, hiking, and kayaking on the Bill Williams River. The habitats of the refuge are a unique blend of Mojave Desert and Sonoran Desert uplands, desert riparian zones, and marsh wetlands habitats, provides for a diverse array of flora and fauna.
Butterflies at Bill Williams River National Wildlife Refuge (NWR) in Arizona. Published by the U.S. Fish & Wildlife Service (USFWS).
Bill Williams River NWR
https://www.fws.gov/refuge/bill_williams_river
https://en.wikipedia.org/wiki/Bill_Williams_River_National_Wildlife_Refuge
The Bill Williams River National Wildlife Refuge protects the lower course of the Bill Williams River, to its mouth at Lake Havasu reservoir, in western Arizona. It is located within eastern La Paz and Mohave Counties, in the Lower Colorado River Valley region. Recreation activities include nature walks, bird watching, hiking, and kayaking on the Bill Williams River. The habitats of the refuge are a unique blend of Mojave Desert and Sonoran Desert uplands, desert riparian zones, and marsh wetlands habitats, provides for a diverse array of flora and fauna.
U.S. Fish & Wildlife Service
Mammals
Bill Williams River
National Wildlife Refuge
Order Rodentia (Rodents)
Family Castoridae
Beaver (Castor canadensis)
Family Erethizontidae
Porcupine (Erethizon dorsaturn)
Desert Bighorn Sheep.
Mammals of the Desert
In the vast desert environments
numerous mammalian species are found.
A drive over the long rough roads on
the refuges gives one the impression
that the desert is devoid of animal
life, but a closer examination reveals
numerous burrows between scattered
bushes, among rocks, and even on the
open plains. The burrows are the home
of ground squirrels, pocket mice, and
kangaroo rats. The home of the familiar
woodrat consist of piles of sticks and
cactus joints that are scattered beneath
bushes and in rock clefts and caves
throughout these refuges.
Desert mammals have adapted their
lives to the extreme temperature and
low humidity of their environments
Water conservation is an absolute
necessity in their activities, The majority
of mammals living in the desert are
nocturnal, foraging only at night when
the relative humidity is higher and
moisture loss from their bodies is
kept at a minimum. Most of the desert
mammals, especially the smaller ones,
have adapted to survive with minimal
water and receive needed moisture from
plant material. During hot summer
days, bighorn sheep lay in the shade of
mountain caves. Large eared mule deer
forage along desert washes at night and
rest during midday in the shade of desert
trees and overhanging banks.
Bats, the only true flying mammals,
find caves, crevices, and mine tunnels
ideal places to congregate during the
day. Most bats in this area are nocturnal
and are rarely seen in the daylight. In
the dim, flickering light of the campfire,
bats may be seen through the night air
catching their meals of insects.
Fanily Geomyidae
Valley Pocket Gopher (Thomomys bottae)
Family Heteromyidae
Merriam’s Kangaroo Rat
(Dipodomys merriami merriami)
Desert Kangaroo Rat
(Dipodomys deserti arizonae)
Arizona Pocket Mouse
(Perognathus amplus)
Little Pocket Mouse
(Perognathus longimembris)
Bailey’s Pocket Mouse
(Perognathus baileyi)
Rock Pocket Mouse
(Perognathus intermedius)
Desert Pocket Mouse
(Perognathus pencillatus pricei)
Spiny Pocket Mouse
(Perognathus spinatus)
Black-tailed Jackrabbitt.
Canyon Mouse
(Peromyscus crinitus disparillis)
Deer Mouse
(Peromyscus maniculatus)
Western Harvest Mouse
(Reithrodontomys megalotis)
Southern Grasshopper Mouse
(Onychomys torridus torridus)
House Mouse (Mus musculus)
Order Lagomorpha (Rabbits and hares)
Family Leporidae
Desert Cottontail
(Sylvilagus audobonii arizonae)
Black-tailed Jackrabbit
(Lepus californicus eremicus)
Family Sciuridae
Occasionally
seen in the upland desert,
Yuma Antelope Squirrel
the
jackrabbit
is not as common as the
(Ammospermophilus harrisii)
cottontail. Jackrabbits may be seen
anytime during the day. On hot summer
Rock Squirrel
(Spermophilus varigatus grammurus) days, they are less active and tend to
rest in the shade. Their young are called
‘leverets’, and are born fully furred, and
Round-tailed Ground Squirrel
(Spermophilus tereticaudus neglectus) with their eyes open. They can outrun
within an hour of birth!
Family Muroidea
Order Insectivora (Shrews and moles)
Muskrat (Ondatra zibethicus)
Family Soricidae
Desert Shrew (Notiosorex crawfordi)
White-throated Woodrat
(Neotoma albigula mearnsi)
Desert Woodrat
(Neotoma lepida auripilla)
Order Chiroptera (Bats)
Family Vespertilionidae
Western Pipistrelle
(Pipistrellus hesperus)
Cactus Mouse
(Peromyscus eremicus eremicus)
Big Brown Bat (Eptesicus fuscus)
Family Antrozoidae
Pallid Bat (Antrozous pallidus)
Family Phyllostomidae
California Leaf-nosed Bat
(Macrotus californicus)
This desert bat possesses a leaf-like,
triangular flap of thick skin projecting
upward from the tip of its nose, thus the
name “leaf nosed.” It roosts in colonies
inside caves and old mine tunnels. They
glean large insects from vegetation such
as sphinx moths and katydids. California
leaf-nosed bats do not hibernate like
many bats, nor do they migrate.
Family Bovidae
Bighorn Sheep (Ovis canadensis
nelsoni) (Ovis canadensis mexicana)
This magnificent animal is most often
seen in dry periods on riverside rocky
escarpments. They are occasionally seen
along the cliffs of the Bill Williams River
where the 2 sub-species are in contact.
Bighorns inhabit the rugged mountain
terrain and slopes which are sparsely
populated with trees, shrubs, and brush.
This brown to grayish brown sheep is
superbly camouflaged in its mountain
habitat and when standing still, the
creamy white rump and white muzzle
Order Carnivora (Carnivores)
Family Felidae
Bobcat (Lynx rufus baileyi)
Family Cervidae
Rocky Mountain Elk (Cervis elaphus)
Mountain Lion (Felis concolor)
California Leaf-nosed Bat.
Family Canidae
Coyote (Canis latrans mearnsi)
Western Red Bat (Lasiurus borealis)
Kit Fox (Vulpes macrotis macrotis)
Hoary Bat (Lasiurus cinereus)
Gray Fox (Urocyon cinereoargenteus)
U.S. Fish & Wildlife Service
Birds
Bill Williams River
National Wildlife Refuge
Birding at Bill Williams River National
Wildlife Refuge
Created in 1941 as part of Havasu
National Wildlife Refuge, the Bill
Williams River National Wildlife
Refuge encompasses approximately 6,000
acres along the Bill Williams and Lower
Colorado Rivers. The four major areas
of the refuge include the open waters
of an arm of Lake Havasu, the Delta
Marsh, riparian woodlands along the Bill
Williams River, and surrounding upland
deserts with dramatic cliffs. The refuge
contains the largest remaining nativevegetation dominated riparian forest
along the Lower Colorado River and the
only one that is still flood re-generated.
Due to the location of the refuge and its
rich habitat diversity, it is an important
breeding, wintering, and migratory
stopover point for birds, as well as
providing habitat for many other
wildlife species.
The 353 bird species are listed in
accordance with the 7th edition A.O.U.
checklist and the 2012 supplement. If you
should find an unlisted or rare species
during your visit, please contact the
Refuge staff and provide a description on
the form on the last page.
Seasonal status and abundance
categories are to be considered as
guides and should not be viewed as
hard seasonal dates because birds and
migration are dynamic and do not follow
the calendar definitions of spring and fall.
Summer status indicated for many nonbreeding species of shorebirds, terns,
flycatchers, vireos, swallows, warblers,
tanagers, and grosbeaks actually refers
to early fall migration, which begins in
July and August for these groups.
As landscapes change due to human
activities and natural processes, species
populations and distributions must
change as well. Over time, species may
become locally absent (extirpated),
restored, newly arrived, endangered,
or even extinct. Some have always been
rare on the Lower Colorado River. On the
Bill Williams River NWR, species once
common but now absent or rare include
Vermilion flycatcher.
wood storks, wood ducks, and vermillion
flycatchers. Species that have recently
arrived include Eurasian collared doves,
indigo buntings, and northern cardinals.
Many riparian obligate species that have
declined in most areas of the southwest
due to habitat changes remain common
on the Bill Williams River where their
habitat has been restored. We manage
habitat to help recover endangered
species as well as declining species
including many riparian forest birds
such as the yellow-billed cuckoo and
summer tanager.
Data for this list are based on Rosenberg,
K.V., R.D. Ohmart. W.C. Hunter and
B.W. Anderson, 1991, Birds of the Lower
Colorado River. University of Arizona
Press; Corman, T. E. and C. Wise-Gervais
(eds), Arizona Breeding Bird Atlas,
2005. University New Mexico Press.,
Audubon Christmas Bird Count http://
birds.audubon.org/christmas-bird-count;
ebird http://ebird.org/content/ebird/; and
refuge records.
How to use your checklist
Symbols used in this list are as follows:
Occurrence
Sp - Spring - period of spring migration
(depending on species) from mid-
February to early June.
S - Summer - June to August.
F - Fall - period of fall migration
(depending on species) from
mid-July to November.
W - Winter - December to February.
Abundance
A - Abundant. A very numerous
species.
C - Common. Certain to be seen in
suitable habitats.
U - Uncommon. Present, not
certain to be seen.
O - Occasional. Seen few times
during a season.
R - Rare. Seen at intervals of
2 to 5 years.
X - Accidental. Fewer than 3 records;
vagrants outside their usual range.
* Confirmed nester.
** Possible nester.
Italicized bird names indicate threatened
or endangered species.
Sp S F W
Ducks, Geese and Swans
Gr. White-fronted Goose u r
Snow Goose u u
Ross’s Goose r r
Brant
x
Canada Goose
u u u
Trumpeter Swan x
Tundra Swan x x
Wood Duck (*historical) r r r r
Gadwall
u x u u
Eurasian Wigeon x
American Wigeon
u x c u
Mallard *
c c c c
Blue-winged Teal
u o u o
Cinnamon Teal *
u o u o
Northern Shoveler
u o u u
Northern Pintail
u o u u
Green-winged Teal
u o u u
Canvasback
u u c
Redhead
u r u u
Ring-necked Duck
c c c
Greater Scaup
c u c
Lesser Scaup
u u u
Surf Scoter
o o o
White-winged Scoter
o r r o
Black Scoter
x x x
Long-tailed Duck
x x x
Bufflehead
u u c
Common Goldeneye
c c c
Barrow’s Goldeneye
u u u
Hooded Merganser r r
Common Merganser
c u c
Red-breasted Merganser u u u
Ruddy Duck
c u c c
New World Quail
Gambel’s Quail *
c c c c
Loons
Red-throated Loon
x x x
Pacific Loon
o x o o
Common Loon
u o u
Yellow-billed Loon
x x x
Western grebe with chick.
Sp S F W
Grebes
Pied-billed Grebe *
c c c c
Horned Grebe
o o u
Red-necked Grebe
x x
Eared Grebe
u o u u
Western Grebe *
c c c c
Clark’s Grebe *
c c c c
Storks
Wood Stork (historical) c c c c
Frigatebirds
Magnificent Frigatebird x x
Boobys
Blue-footed Booby x
Brown Booby x
Cormorants
U.S. Fish & Wildlife Service
Butterflies
Bill Williams River
National Wildlife Refuge
Butterflies at Bill Williams River NWR
The Bill Williams River National Wildlife Refuge supports the last and largest
remnant of the native riparian habitats
that once filled the valleys of the lower
Colorado River and its tributaries. These
now unique habitats support many butterfly species. A total of 34 species have
been documented in the refuge, 11 (32%)
of which were found only here in a recent
study (Nelson and Andersen 1998). One
species, MacNeill’s sootywing skipper, is
of some concern to conservationists due
to its rarity. The larvae apparently only
feed on one species of saltbush (Atriplex lentiformis), a native shrub of the
lower Colorado River valley. Wiesenborn
(1997) often observed adults flying between the saltbush and honey mesquite
(Prosopis glandulosis).
Scientific name
Common name
Swallowtails (Papilionidae)
Papilio polyxenes
Black Swallowtail
Papilio Zelacaon
Zelacaon Swallowtail
Whites and Sulphurs (Pieridae)
Pontia protodice
Checkered White
Colias eurytheme
Orange (Alfalfa) Sulphur
Phoebis sennae
Cloudless Sulphur
Eureme nicippe
Sleepy Orange (Nicippe Yellow)
Nathalis iole
Dainty Sulphur
Great Purple Hairstreak
Gossamer-winged Butterflies
(Lycaenidae)
Atlides halesus
Great Purple Hairstreak
Strymon melinus
Gray Hairstreak
Metalmarks (Riodinidae)
Calephelis nemesis
Fatal Metalmark
Ministrymon leda
Leda Ministreak
Apodemia mormo
Mormon Metalmark
Brephidium exile
Pygmy Blue
Apodemia palmeri
Palmer’s Metalmark
Leptotes marina
Marine Blue
Brush-footed Butterflies (Nymphalidae)
Libytheana carinenta
Snout Butterfly
Hemiargus ceraunus
Cerannus Blue
Hemiargus isola
Reakirt’s Blue
Calephelis wrighti
Wright’s Metalmark
Euptoieta claudia
Variegated Fritillary
Nymphalis antiopa
Mourning Cloak
Vanessa (Cynthia) cardui
Painted Lady
Vanessa atalanta
Red Admiral
References
Levy, J. N. 1996. Pers.comm.
Milne, L and M. Milne. 1980. Audubon
Society Field Guide to North American
Insects and Spiders. Alfred Knopf, N.Y.
988 pgs.
Nelson, S. M. and D. C. Anderson.
1998. Butterfly (Papilionoidea and
Hesperioidae) communities associated
with some natural and altered riparian
habitats along the lower Colorado River.
Technical Memorandum No. 8220-98-3.
32 pgs.
Tilden, J. W. and A. C. Smith. 1986. A
Field Guide to Western Butterflies.
Houghton Mifflin, Boston. 370 pgs.
Wiesenborn, W. D. 1997. Hesperopsis
gracielae (MacNeill) (Lepidoptera:
Hesperiidae) flight between hostplants
and Prosopis glandulosa Torrey. PanPacific Entomologist 73(3):186-189.
Common buckeye.
Jumonia coenia
Common Buckeye
Hylephila phyleus
Fiery Skipper
Limenitis arthemis
Red Spotted Purple
Lerodea eufala
Euphala Skipper
Limenitis (Basilarchia) archippus
Viceroy
Danaus plexippus
Monarch
Danaus gilippus
Queen
Skippers (Hesperiidae)
Erynnis funeralis
Funereal Duskywing
Pyrugus scriptura
Small Checkered Skipper
Pyrgus communis (Grote)
Common Checkered Skipper
Heliopetes ericetorum
Northern White-Skipper
Hesperopsis gracielae (MacNiell)
MacNiell’s Sootywing Skipper
Copaedes aurantiacus
Orange Skipperling
Viceroy.